Arthur and Donald are the twins of Helena (played by Tatiana Maslany in the original Orphan Black). Their biological father was Henrik Johanssen (Peter Outerbridge), a religious cult leader, who impregnated Helena and other women without their consent using artificial insemination. Helena killed him for it, but chose to keep the children and raise them on her own. Having been raised as an assassin in Ukraine, Helena used her skills to teach Arthur and Donald how to survive in the wild since they were kids. By the time of Orphan Black: Echoes, they would be about 38 years old.
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