Story Plot

A zebra named Marty lived his entire life at the Central Park Zoo with his best friends since childhood: a lion named Alex, who is Marty's best friend, a giraffe named Melman and a hippopotamus named Gloria. Alex enjoys showing off for the public and his celebrity status as "the king of New York". Marty, on the other hand, has grown bored with his daily routine and longs to experience the wild. On Marty's tenth birthday, Alex, Melman and Gloria attempt to cheer him up, but Marty, still unsatisfied, learns that the zoo's penguins—Skipper, Kowalski, Rico, and Private—are trying to escape, and follows them out. Alex, Melman and Gloria pursue Marty and attempt to convince him to return. The four, along with the penguins and two chimpanzees named Mason and Phil, converge at Grand Central Station where the authorities sedate them using tranquillizer guns. Under pressure from anti-captivity activists, the zoo is forced to ship the escaped animals by sea to a Kenyan wildlife preserve. During their travels, the penguins escape and take over the ship, intent on taking it to Antarctica. Their antics on the bridge cause the crates containing Alex, Marty, Melman, and Gloria to fall overboard and wash ashore on Madagascar.

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